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軟骨魚綱

軟骨魚綱
屬 種 發現地點 發現部位 備註 圖片
無尖齒魚科未定屬
Acrodontidae indet.[1] 		未定種 屬於弓鮫目
Bahariyodon[1] B. bartheli 屬於弓鮫目
Cenocarcharias[1] C. tenuiplicatus 一顆牙齒[1] 屬於白堊刺甲鯊科
Distobatus[1] D. nutiae 屬於弓鮫目
哈氏鯊科（英語：Haimirichia）
Haimirichia[1] 		H. amonensis 一顆牙齒[1] 屬於鼠鯊目
Marckgrafia[1] M. lybica 13顆牙齒[1] 屬於鰩總目
帆鋸鰩屬
Onchopristis 		努米底亞帆鋸鰩
O. numida[2] 		目前已知體型最大的鋸鰩，屬於鰩目[3]
 
帆鋸鰩的吻突與牙齒化石
Peyeria[1] P. libyca 3顆牙齒[1] 可能為努米底亞帆鋸鰩的次異名
外側棘鯊屬（英語：Tribodus）
Tribodus[1] 		T. sp. 屬於弓鮫目

條鰭魚總綱

條鰭魚總綱
屬 種 發現地點 發現部位 備註 圖片
阿德里亞娜魚屬
Adrianaichthys[1] 		A. pankowskii 鱗片[4]與兩具顱骨[5] 屬於雀鱔目，原本被認為屬於鱗齒魚屬（英語：Lepidotes），但之後被重新分類至獨立屬[6]
 
埃達克龍魚屬（英語：Aidachar）復原圖
Afrocascudo（英語：Afrocascudo）[7] A. saharaensis 屬於新鰭亞綱，可能屬於甲鯰科或屬於雀鱔目奧巴魚科的幼年個體[8]
Agassizilia（英語：Agassizilia）[9] A. erfoudina 可能屬於堅齒魚科
阿古爾特魚屬
Agoultichthys[1] 		A. chattertoni 屬於條鰭魚總綱，但詳細分類位置不明，可能屬於巨半椎魚科（英語：Macrosemiidae）[10]或Ophiopsiellidae[11]
埃達克龍魚屬（英語：Aidachar）
Aidachar 		A. pankowskii 屬於乞丐魚目
巴奇魚屬
Bartschichthys[1] 		B. sp. 背鰭的的刺狀鰭骨[1] 屬於腕鰭魚綱
巴氏巴威多鰭魚
Bawitius 		cf. B. sp. 鱗片與下顎碎片[4] 屬於腕鰭魚綱
卡拉矛普萊弓鰭魚屬（英語：Calamopleurus）
Calamopleurus[1] 		非洲卡拉矛普萊弓鰭魚
C. africanus 		部分顱骨[1] 屬於弓鰭魚目
Concavotectum（英語：Concavotectum）[1] C. moroccensis 屬於塞納命運魚目
齒雀鱔屬
Dentilepisosteus[1] 		D. kemkemensis 屬於雀鱔目
雙頜鯡屬（英語：Diplomystus）
Diplomystus[1] 		D. sp. 屬於骨鯡群真骨類
Diplospondichthys[1] D. moreaui A member of Actinopterygii of uncertain phylogenetic placement, possibly a teleost
Erfoudichthys[1] E. rosae Isolated skull[1] A small-bodied teleost of unknown affinity
Neoproscinetes[9] N. africanus A member of the family Pycnodontidae
Obaichthys O. africanus Isolated scales[4] A member of Lepisosteiformes
Oniichthys O. falipoui Near complete skeleton including skull[4] A member of Lepisosteiformes
Palaeonotopterus[1] P. greenwoodi A member of Osteoglossomorpha
Serenoichthys[1] S. kemkemensis Several articulated skeletons[1] A small cladistian
Spinocaudichthys[1] S. oumtkoutensis An elongate freshwater acanthomorph
Stromerichthys S. aethiopicus
Sudania[1] S. sp. An isolated pinnula[1] A cladistian

肉鰭魚總綱

肉鰭魚總綱
屬 種 發現地點 發現部位 備註 圖片
Arganodus A. tiguidiensis A lungfish
 
Life restoration of Axelrodichthys
 
The Queensland Lungfish, the only living member of Neoceratodus
Axelrodichthys[12] A.? lavocati A mawsoniid coelacanth; this species was previously assigned to Mawsonia, and its generic assignment is still not certain[13]
Neoceratodus N. africanus A lungfish

Amphibians

Amphibians
屬 種 發現地點 發現部位 備註 圖片
Anura indet.[14] Indeterminate Douira Formation Incomplete left ilium
Cretadhefdaa[14] C. taouzensis Douira Formation Posterior portion of the skull, incomplete squamosal, incomplete maxilla, three incomplete presacral vertebrae, one incomplete sacral vertebra A neobatrachian frog with possible hyloid affinities.
cf. Kababisha[15] Indeterminate A salamander belonging to the family Sirenidae
?Neobatrachia indet.[14] Indeterminate Douira Formation Incomplete humerus A frog, possibly a member of Ranoidea.
Oumtkoutia[15] O. anae A frog belonging to the family Pipidae

Lizards and snakes

色鍵
分類單元 重新分類的分類單元 錯誤的分類單元 可疑的分類單元或初級同義詞 足跡分類單元 蛋分類單元 形態分類群
註釋
不確定或暫定的分類單元以小字體展示； 被劃掉的分類單元不可信
Lizards and snakes reported from the Continental Red Beds
屬 種 發現地點 發現部位 備註 圖片

Bicuspidon

B. hogreli[16]

A polyglyphanodontid lizard.

Jeddaherdan[17]

J. aleadonta

Partial mandible with teeth.

An iguanian belonging to the group Acrodonta, possibly a relative of the uromastycine agamids. Argued by Vullo et al. (2022) to actually come from Quaternary beds, and to be based on a fossil material of a member of the genus Uromastyx.[18]

Lapparentophis

L. ragei[19]

Two isolated trunk vertebrae

An early snake.

Madtsoiidae indet.[15]

Indeterminate

Vertebrae[1]

An early snake.

?Nigerophiidae indet.[15]

Indeterminate

Dorsal vertebrae[1]

An early snake.

Norisophis[20]

N. begaa[20]

One posterior and two mid-trunk vertebrae

A stem-snake.

Indeterminate[20]

A mid-trunk vertebra

Simoliophis[15]

cf. S. libycus

Vertebrae[1]

An early snake.

Plesiosaurs

Plesiosaurs
屬 種 發現地點 發現部位 備註 圖片
Leptocleididae cf. Leptocleidus[21] indeterminate

Turtles

Turtles reported from the Continental Red Beds
屬 種 發現地點 發現部位 備註 圖片

Dirqadim

D. schaefferi

A Euraxemydid

Galianemys

G. emringeri

A Cearachelyin

G. whitei

Hamadachelys

H. escuilliei

Crocodylomorphs

A tooth enamel identified as cf. Sarcosuchus was discovered from the Ifezouane Formation.[22]

Crocodylomorphs reported from the Continental Red Beds
屬 種 發現地點 發現部位 備註 圖片

Aegisuchus

A. witmeri

"Partial braincase of a large individual with skull roof, temporal, and occipital regions."[23]

An aegyptosuchid that may be a synonym of Laganosuchus.[1]

 
Aegisuchus
 
Araripesuchus
 
Elosuchus cherifiensis
 
Laganosuchus

Antaeusuchus[24]

A. taouzensis

Paired mandibles and a partial right mandible

A peirosaurid.

Araripesuchus

A. rattoides

Douira Formation

Elosuchus

E. cherifiensis

  • Gara Sbaa Formation
  • Douira Formation

An elosuchid. The material may represent two different species.[1]

Hamadasuchus

H. rebouli

  • Gara Sbaa Formation
  • Douira Formation?

A peirosaurid.

Kemkemia

K. auditorei

Errachidia Province, Morocco[25]

Known from an isolated caudal vertebra.[25]

Initially thought to be a neotheropod,[25] but subsequently discovered to be an indeterminate crocodyliform.[26]

Laganosuchus

L. thaumastos

A stomatosuchid.

Lavocatchampsa[27]

Lavocatchampsa sigogneaurusselae

Anterior portion of a rostrum with mandible, with an almost complete dentition[27]

A candidodontid notosuchian.[27]

Dinosaurs

Indeterminate lithostrotian remains once misattributed to the Titanosauridae are present in the province of Ksar-es-Souk, Morocco.[28]

色鍵
分類單元 重新分類的分類單元 錯誤的分類單元 可疑的分類單元或初級同義詞 足跡分類單元 蛋分類單元 形態分類群
註釋
不確定或暫定的分類單元以小字體展示； 被劃掉的分類單元不可信

Ornithischians

Ornithischians reported from the Continental Red Beds
屬 種 發現地點 發現部位 備註 圖片

Ankylosauria[1][29]

Indeterminate

Douira Formation

An isolated tooth.[1]

A probable ankylosaur[29]

Ornithopoda[30]

Indeterminate

Douira Formation

A large, clover-shaped, three-toed footprint.[1]

Comparable in size and shape to tracks typically attributed to Iguanodon.[30]

Sauropods

Sauropods reported from the Continental Red Beds
屬 種 發現地點 發現部位 備註 圖片

Rebbachisaurus

R. garasbae

Ksar-es-Souk province, Morocco.[28]

Gara Sbaa Formation

A rebbachisaurid.

Somphospondyli[31]

Indeterminate

Anterior dorsal vertebra, partial right ischium[31]

The vertebra might belong to a basal titanosaurian, possibly distinct from Aegyptosaurus and Paralititan.[31] The ischium is not identifiable beyond Somphospondyli; it preserves numerous grooves and pits which might be feeding traces left by a very large non-avian theropod.[31]

Titanosauria[32][1]

Indeterminate

  • Ifezouane Formation
  • Douira Formation

Isolated teeth, caudal vertebrae, a partial humerus, a tarsal bone and the proximal end of an ulna.[1]

Fossil material of one or more titanosaurian sauropods. Some fossils are indicative of large body size comparable to Paralititan stromeri.[1]

Theropods

Theropods reported from the Continental Red Beds
屬 種 發現地點 發現部位 備註 圖片

Abelisauridae[33]

Indeterminate

Isolated teeth.[33][34]
Partial right femur.[35] Partial right maxilla.[1] A cervical vertebra.[36] Left maxilla.[1]

Abelisaurid material belonging to one or two distinct taxa.[1]

 
Carcharodontosaurus saharicus
 
Deltadromeus agilis
 
Rebbachisaurus garasbae
 
Rugops primus
 
Spinosaurus aegyptiacus reconstruction

Carcharodontosaurus[28]

C. saharicus[28]

Ksar-es-Souk province, Morocco.[28] Douira Formation Partial skull, including braincase, nasals, postorbitals, jugals, left lacrimal and right maxilla with most teeth.[37]

A carcharodontosaurid theropod.

Carcharodontosauridae[38] Indeterminate Southeast of Taouz, Errachidia Province Ifezouane Formation partial maxilla and partial jugal A carcharodontosaurid theropod different from C. saharicus

Deltadromeus

D. agilis

Gara Sbaa Formation

"Partial skeleton, isolated limb elements."[39]

A noasaurid ceratosaurian or possible neovenatorid carnosaur. May be synonymous with Bahariasaurus.

Dromaeosauridae[33]

Indeterminate

Isolated teeth.[33]

Originally described as teeth of indeterminate dromaeosaurids. Hendrickx et al. (2024) reinterpreted this fossil material as teeth of abelisauroid theropods, including noasaurids and juvenile abelisaurids.[34]

cf. Elaphrosaurus

Indeterminate

Ksar-es-Souk province, Morocco.[28]

Fossils previously referred to cf. Elaphrosaurus are actually indeterminate theropod remains.

Noasauridae[36]

Indeterminate

An anterior cervical vertebra[36]
Isolated teeth[34]

"Osteoporosia"[40] "O. gigantea"[40] A tooth and a possible neural arch from another specimen.[40] A theropod, possibly synonymous with Sauroniops.[41]

Saurischia[42]

Indeterminate

An isolated cervical vertebra.[42]

An indeterminate saurischian.

Sauroniops[43]

S. pachytholus

Ifezouane Formation

"An isolated and almost complete left frontal,[44] and a possible tooth and neural arch from two other specimens."[41]

A carcharodontosaurid distinct from Carcharodontosaurus.[43][44]

Spinosaurus

S. aegyptiacus

Ksar-es-Souk province, Morocco.[28] Douira Formation Partial skeleton, including parts of the skull, neck, torso, and most of the tail and hind limbs. [45]

Numerous isolated bones.

Pterosaurs

Pterosaurs of the Kem Kem Beds
屬 種 發現地點 發現部位 備註 圖片

Afrotapejara[46]

A. zouhri[46]

Takmout Ifezouane Formation

A fragment of bone interpreted as a fragment of anterior mandibular symphysis,[47] and additional jaw fragments that pertain to the rostrum as well as indeterminate jaw fragments.[48]

A tapejarid pterosaur. Originally believed to belong to either the family Thalassodromidae[49] or an additional specimen of Alanqa saharica.[50]

 
Afrotapejara zouhri
 
Akharhynchus martilli
 
Alanqa saharica
 
Anhanguera
 
Coloborhynchus
 
Leptostomia
 
Ornithocheirus
Akharhynchus[51] A. martilli Tafilalt Ifezouane Formation A fragment of the anterior part of the premaxillae A tropeognathine anhanguerian.

Alanqa[49]

A. saharica[49]

Ifezouane Formation The holotype is a mandibular symphysis, of different parts of the jaw A pterosaur of uncertain phylogenetic placement, probably an azhdarchid.[48]

Anhanguera[52]

A. cf. piscator[52]

upper Ifezouane Formation

Partial mandibular symphysis[52]

Apatorhamphus[53]

A. gyrostega[53]

Ifezouane Formation

Partial rostrum and mandible, with additional referred jaw fragments[48]

A possible chaoyangopterid azhdarchoid pterosaur.[53] Originally believed to be a possible pteranodontid,[49] a possible dsungaripterid,[54] a possible non-azhdarchid azhdarchoid or nyctosaurid,[54] or a specimen of Alanqa saharica.[50]

Azhdarchidae indet.[54]

Indeterminate[54]

Three middle cervical vertebrae.[54][49]

Averianov (2014) considered these vertebrae to pertain to Alanqa saharica,[50] although the vertebrae may be indicative of two taxa.[54]

Coloborhynchus[52] C. sp. A.[52] Hassi El Begaa Premaxillae fragment[52] Possibly a specimen of Nicorhynchus fluviferox.[55]

Leptostomia[56]

L. begaaensis[56]

Aferdou N』 Chaft

upper Ifezouane Formation

Partial rostrum and partial mandibular synthesis[56]

A small, long-beaked pterosaur, likely a member of Azhdarchoidea.[56]

Nicorhynchus[55]

N. fluviferox[57][55]

Possibly Aferdou N』Chaft, Hassi El Begaa[55]

Ifezouane Formation

An anterior portion of the rostrum.[55]

Originally described as a species of Coloborhynchus[57] but subsequently transferred to the genus Nicorhynchus.

Ornithocheirus[52]

O. cf. simus.[52]

upper Ifezouane Formation

Premaxillae fragment[52]

Siroccopteryx[58]

S. moroccensis[58]

Anterior part of a rostrum

Classified by some authors as a species belonging to the genus Coloborhynchus.[49]

Xericeps X. curvirostra Aferdou N'Chaft Ifezouane Formation Mandibular symphysis and partial mandible[48] An indeterminate azhdarchoid, possibly a chaoyangopterid.[48]
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